Land mollusk surveys on USFS Northern Region lands

Land Mollusk Surveys on
USFS Northern Region Lands

Prepared for:

USDA Forest Service, Northern Region

P.O. Box 7669

Missoula, MT 59807

By:
Paul Hendricks, Bryce A. Maxell and Susan Lenard

Montana Natural Heritage Program

Natural Resource Information System

Montana State Library

May 2006

MONTANA

Natural Heritage
Program

Land Mollusk Surveys on
USFS Northern Region Lands

Prepared for:

USDA Forest Service, Northern Region

P.O. Box 7669

Missoula, MT 59807

Agreement Number:
#05-CS-110115600-033

By:
Paul Hendricks, Bryce A. Maxell and Susan Lenard

JW*" MONTANA

yft Natural Heritage
^Program

fe T State It |2& Natural Resource

/> Library V^0 Information System

© 2006 Montana Natural Heritage Program

P.O. Box 201800 • 1515 East Sixth Avenue • Helena, MT 59620-1800 • 406-444-5354

This document should be cited as follows:

Hendricks, P., B. A. Maxell and S. Lenard. 2006. Land Mollusk Surveys on USFS Northern Region
Lands. A report to the USDA Forest Service, Northern Region. Montana Natural Heritage Program,
Helena, Montana. 11 pp. plus appendices.

Executive Summary

Using prior published reports and the NatureServe
web site as starting points, we compiled a list of 29
snail taxa within the U.S. Forest Service Northern
Region (Region 1) area ranked in 2005 as G1-G3
or T1-T3, thereby meeting USFS Species of
Concern (SOC) criteria, and two additional G5 snail
taxa ranked S1-S2, thereby meeting USFS Species
of Interest (SOI) criteria. We also compiled a list
of eight slug taxa ranked in 2005 as G1-G3, and
three additional slug taxa ranked G4-G5 but S 1-S2,
again meeting the respective USFS criteria for
SOC or SOI.

We conducted a total of 108 site surveys in 2005
targeting areas lacking previous information on
terrestrial mollusks. These were distributed on the
Montana Forests as follows: Beaverhead-
Deerlodge (13), Bitterroot (0), Custer (8), Flathead
(9), Gallatin (17), Helena (13), Kootenai (5), Lewis
& Clark (19), Lolo (1). We distributed surveys on
the Idaho Forests as follows: Clearwater (11),
Idaho Panhandle (4), Nez Perce (8).

We documented 83 new locations for ten SOC taxa
and three SOI taxa during our 2005 surveys:
Selway Forestsnail Allogona lombardii (3 sites),
Nimapuna Tigersnail Anguispira nimapuna (4
sites), Striate Disc Discus shimeki (1 site), Berry's
Mountainsnail Oreohelix strigosa berryi (6 sites),
Robust Lancetooth Haplotrema vancouverense (4
sites), Humped Coin Polygyrella polygyrella (6
sites), Fir Pinwheel Radiodiscus abietum (16
sites), Pale Jumping-slug Hemphillia camelus (9
sites), Pygmy Slug Kootenaia burkei (6 sites),
Reticulate Taildropper Prophysaon andersoni (1
site), Smoky Taildropper Prophysaon humile (16
sites), Lyre Mantleslug Udosarx lyrata (2 sites),

and Sheathed Slug Zacoleus idahoensis (9 sites).
Most locations are from west of the Continental
Divide in mesic forest habitats (e.g., western
redcedar, western hemlock, mesic Douglas-fir,
grand fir).

Two SOC and one SOI slug species, all discovered
on the Kootenai National Forest, are new to the
known mollusk fauna of Montana: Pale Jumping-
slug, Pygmy Slug, and Reticulate Taildropper. The
2005 survey added seven new Montana locations
and nine new Idaho locations for a third SOC slug
species, Smoky Taildropper, which was
documented in Montana only once prior to 2004.
As a result of the 2005 surveys, Global Ranks
shifted downward for five species (Humped Coin,
Fir Pinwheel, Pale Jumping-slug, Pygmy Slug, and
Smoky Taildropper). We also collected distribution
data on 21 additional non-SOC species as we
encountered them during our surveys.

At least some SOI G4-G5 taxa found during our
2005 surveys may prove to be distinct from related
coastal populations, as their disjunct distributions
are similar to some vertebrate amphibian taxa (e.g.,
Dicamptodon, Ascaphus, Plethodon) now split
into coastal and Rocky Mountain sister species.
Therefore, we think it desirable to conduct genetic
analyses of several mollusk SOC and SOI taxa to
determine it they represent forms that merit full
species status. Additional inventory is also
desirable to fill remaining distribution gaps, describe
habitat associations thoroughly enough to develop
predictive distribution models, and lay the
foundation for developing a long-term monitoring
scheme and standardized survey methodology.

Acknowledgements

Fred Samson (USFS) recognized the need to
address invertebrates in the Forest planning
process, appreciating the extremely limited
information available for management decision-
making, and promoted the project through the
USFS Regional Inventory and Monitoring (RIM)
program. Henning Stabins (Plum Creek Timber
Company) and the Amphibian Inventory Project
provided us with additional records of SOC mollusk
species that helped fill significant gaps in
distributions. Bill Bosworth, zoologist with the
Idaho Conservation Data Center, provided the
Montana Natural Heritage Program (MTNHP)

with location data on SOC species tracked in the
Idaho portions of the Northern Region; the Idaho
records were especially critical for the production
of new distribution maps, and fleshing out
distributions of rare land mollusk species that occur
on both sides of the Idaho-Montana border. Bill
Leonard (Olympia, WA) and Tim Pearce (Carnegie
Museum of Natural History) verified our tentative
SOC and SOI slug identifications. Matthew Gates
assisted with field surveys. Coburn Currier
(MTNHP) assisted on some surveys and formatted
and printed this report. We thank them all.

IV

Table of Contents

Introduction 1

Methods 2

Results and Discussion 5

Overview 5

Species Accounts 6

Conclusions and Recommendations 9

References Cited 10

Appendix A. Global/State Rank Definitions

Appendix B. USFS Region 1 Survey Sites for Land Mollusks in 2005

Appendix C. Standaridized Data Sheets

Appendix D. Example Species Photograpshs

Appendix E. Distribution Maps for SOC/SOI Land Mollusks on USFS Region 1 Lands

Appendix F. Distribution Maps for Non-SOC Land Mollusks on USFS Region 1 Lands

Appendix G. Distribution Maps for Exotic Land Mollusks on USFS Region 1 Lands

List of Tables

Table 1. Species of Concern land mollusks: distribution by Forest in Region 1 2

Table 2. Species of Concern land mollusks: habitat associations 3

Table 3. Number of survey sites where Species of Concern land mollusks were detected on Northern

Region Forests during 2005 survey 5

Introduction

Within and adjacent to the landscape bounded by
the Northern Region are a large number of land
mollusk species that are endemic to the Northern
Rocky Mountains, and several additional species
are restricted to the Pacific Northwest, with
disjunct populations in northern Idaho and
northwestern Montana (Frest and Johannes 1995,
1997, 2001; Hendricks 2003). Parts of the survey
area, especially portions of the Lower Salmon
River drainage, were recognized relatively early as
centers of mollusk endemism, and drew attention of
several early collectors (Frest and Johannes 1997).
Nevertheless, many areas in Idaho as well as
Montana were never visited or remain poorly
inventoried, as demonstrated by the recent
discovery of a new slug genus in northern Idaho
(Leonard et al. 2003). Limited survey of the region
is partly a result of timing. When much pioneering
collecting of the terrestrial mollusk fauna was
undertaken, roughly 1860-1950, many portions of
the survey area were difficult to reach without
significant commitment of time and resources.
During the mid and late 20 th century, when road

access across the study area increased
dramatically, far fewer malacologists were resident
or active in the region. Only recently has there
been recognition by biologists that many mollusk
species in the region are threatened with a variety
of potentially detrimental land use activities,
prompting renewed inventories.

The FS is required under the National Forest
Management Act (1976) and Code of Federal
Regulations (CFR 1985) to maintain a diversity of
plant and animal species. Inventory is a first step
in the evaluation of landscapes and their likelihood
of supporting populations of animal species of
conservation concern. Pursuant with this
legislation and associated regulations, the Northern
Region initiated surveys in 2005 for a suite of land
mollusks listed as Species of Concern in Montana
and Idaho. Objectives of the 2005 inventory
included filling species distribution gaps, testing
survey methodology, and collecting geospatial and
habitat data for the development of predictive
habitat models that can aid future survey efforts.

Methods

Prior to conducting field surveys, we searched the
published and gray literature to compile a list of
high-priority "target" species (globally and state
rare species in Montana, and globally rare species
in Idaho). Primary sources we used for this
compilation included Pilsbry (1939, 1948), Frest and
Johannes (1995, 1997, 2001), and Hendricks
(2003). This resulted in a list of 42 species and
subspecies (31 snails, 11 slugs) we considered to be

of conservation concern (Table 1); only eight of
these taxa are ranked less than G3 (see Appendix
A for ranks definitions). We then generated a list
of general habitat associations for the high-priority
species (Table 2), to help us prioritize habitats for
survey. Limited information for mollusks east of
the Continental Divide in Montana made this
process more problematic for the high-priority
species that occur or might occur in that region.

Table 1. Species of Concern land mollusks: distribution by Forest in Region 1. G Ranks of species in bold changed as a
result of our 2005 surveys (see text for previous ranks).

Montana 3

Idaho"

SPECIES | G RANK

B-D | BI | CU | FL | GA | HE | KO | L-C | LO

CL | IP | N-P

Snails

Allogona lombardii (ID)

Gl

X

Allogona ptychophora solida (ID)?

G5T2T3

? !

Anguispira nimapuna (ID)

Gl

X

X

Crypt omastix harfordiana (ID)?

G3G4

?

Cryptomastix magnidentata (ID)?

Gl

?

Crypt omastix mullani blandi (ID)?

G4T1

?

Cryptomastix mullani clappi (ID)

G4T1

X

Cryptomastix sanburni (ID)?

Gl

?

Discus brunsoni (MT)?

Gl

?

Discus marmorensis (ID)

G1G3

X

Discus shimekii (MT, ID?)

G5

X

X

?

Haplotrema vancouverense* (MT, ID)

G5

X

X

X

Helicodiscus salmonaceus (ID)

G1G2

X

Oreohelix alpina (MT)

Gl

X

Oreohelix amariradix (MT)

G1G2

X

Oreohelix carinifera (MT)

Gl

X

Oreohelix elrodi (MT)

Gl

X

Oreohelix hammeri (ID)

Gl

X

Oreohelix idahoensis baileyi (ID)

G1G2T1

X

Oreohelix idahoensis idahoensis (ID)?

G1G2T1T2

?

Oreohelix inter sum (ID)?

Gl

?

Oreohelix jugalis (ID)?

Gl

?

Oreohelix strigosa berryi (MT)

G5T2

X

X

X

X

Oreohelix strigosa goniogyra (ID)

G5T1Q

X

Oreohelix vortex (ID)?

G1G3

?

Oreohelix waltoni (ID)?

G1G3

?

Oreohelix yavapai mariae (MT)

G4T1

X

Planogyra clappi (ID)

G3G4

X

Polygyrella polygyrella (MT, ID)

G3

X

X

X

Pristiloma idahoense (ID)

G2G3

X

Radiodiscus abietum (MT, ID)

G4

X

X

X

X

X

X

X

Slugs

Hemphillia camelus* (MT, ID)

G4

X

X

X

X

X

Hemphillia danielsi (MT)

G2G3

X

?

Kootenaia burkei* (MT, ID)

G2

X

X

Magnipelta mycophaga (MT, ID)

G3

X

X

X

X

X

X

Prophysaon andersoni* (MT, ID)

G5

X

?

Prophysaon coeruleum (ID)

G4

X

Prophysaon dubium (ID)

G4

X

Prophysaon humile* (MT, ID)

G3

X

X

X

X

X

X

Udosarx lyrata lyrata (MT, ID)

G2T2

X

?

X

Udosarx lyrata russelli (MT)?

G2T1

?

Zacoleus idahoensis (MT, ID)

G3G4

X

X

X

X

X

X

a Montana Forests codes: Beaverhead-Deerlodge (B-D), Bitterroot (BI), Custer (CU), Flathead (FL), Gallatin (GA), Helena

(HE), Kootenai (KO), Lewis & Clark (L-C), Lolo (LO).

b Idaho Forest codes: Clearwater (CL), Idaho Panhandle (I-P), Nez Perce (N-P).

* new species for Montana SoC list

? taxon apparently not yet recorded on USFS Region 1 lands, but in area and should be looked for

Table 2. Species of Concern land mollusks: habitat associations. G Ranks of species in bold changed c
result of our 2005 surveys (see text for previous ranks).

Moist Mixed-conifer Forest
Riparian

Aspen

Dry Mixed-conifer Forest

Limestone
Talus*

Species

G Ranks

Cedar-
hemlock,
grand fir,
Doug-fir

Spruce-
fir

Talus-
rocky
ground

Ponderosa

pine,
Doug-fir

Juniper-
sage

Talus-
rocky
ground

SNAILS

Allogona lombardii (ID)

Gl

X

Allogona ptychophora solida (ID)?

G5T2T3

X

X

Anguispira nimapuna (ID)

Gl

X

X

X

X

Cryptomastix harfordiana (ID)?

G3G4

X

X

Cryptomastix magnidentata (ID)?

Gl

X

X

X

Cryptomastix milliard blandi (ID)?

G4T1

X

Cryptomastix mullani clappi (ID)

G4T1

X

Cryptomastix sanburni (ID)?

Gl

X

Discus brunsoni (MT)?

Gl

X

Discus marmorensis (ID)

G1G3

X

X

X

X

Discus shimekii (MT, ID?)

G5

X

X

Haplotrema vancouverense (MT,ID)#

G5

X

Helicodiscus salmonaceus (ID)

G1G2

X

X

Oreohelix alpina (MT)

Gl

X

X

Oreohelix amariradix (MT)

G1G2

X

X

Oreohelix carinifera (MT)

Gl

X

X

X

X

Oreohelix elrodi (MT)

Gl

X

Oreohelix hammeri (ID)

Gl

X

X

Oreohelix idahoensis baileyi (ID)

G1G2T1

X

X

0. i. idahoensis (ID)?

G1G2T1T2

X

X

X

Oreohelix inter sum (ID)?

Gl

X

Oreohelix jugalis (ID)?

G1G2

X

Oreohelix strigosa berryi (MT)

G5T2

X

X

X

0. s. goniogyra (ID)

G5T1Q

X

X

Oreohelix vortex (ID)?

G1G3

X

Oreohelix waltoni (ID)?

G1G3

X

X

Oreohelix yavapai mariae (MT)

G4T1

X

X

Planogyra clappi (ID)

G3G4

X

Polygyrella polygyrella (MT, ID)

G3

X

X

X

X

Pristiloma idahoense (ID)

G2

X

X

Radiodiscus abietum (MT, ID)

G4

X

X

X

SLUGS

Hemphillia camelus (MT, ID)

G4

X

X

Hemphillia danielsi (MT)

G2G3

X

X

Kootenaia burkei (MT, ID)

G2

X

Magnipelta magnipelta (MT, ID)

G3

X

X

X

X

Prophysaon andersoni (MT, K))#

G5

X

Prophysaon coeruleum (ID)#

G4

X

Prophysaon dubium (ID)#

G4

X

X

Prophysaon humile (MT, ID)

G3

X

X

X

Udosarx lyrata lyrata (MT, ID)

G2T2

X

X

£/. /. rw^//* (MT)?

G2T1

X

X

Zacoleus idahoensis (MT, ID)

G3G4

X

X

#• These low G-rank taxa may prove to be distinct from coastal populations, as their disjunct distributions are similar to
individual vertebrate taxa (e.g. Dicamptodon, Ascaphus, Plethodon) now split into coastal and Rocky Mountain species.
* Limestone talus associates may occur in either dry or moist sites, but are most often limestone or limestone-derived soil
obligates

We conducted field surveys for land mollusks
during mid- September to early November 2005,
when the weather was most suitable (cool and
moist) for finding active snails and slugs. We
attempted to visit all 12 National Forests in the
Northern Region, so survey effort was stratified by
Forest (sometimes by mountain range within a
forest) and spread thinly across the inventory area
(sites surveyed are listed in Appendix B).
However, we spent the most time on Forests with

the least amount of prior survey effort or
distribution information. Forests in Idaho were an
exception because we were not aware at the time
of our 2005 surveys how much survey history they
already had.

We selected sites for surveys based primarily on
the presence of perennial water, moist mature
conifer forest, aspen, and/or limestone talus or
other rock outcrops. At each site, we conducted

timed surveys while searching under leaf litter,
dead wood and bark, rocks imbedded in the ground,
or digging into talus. Usually within a survey site
we searched several locations with habitat features
(such as bryophyte mats, dead wood and imbedded
rocks, or talus slopes) considered by experts to be
favored by snails and slugs, often concentrating
searches in riparian zones.

We recorded a variety of habitat and site
information at each survey location on standardized
data forms (Appendix C). Survey data have been

entered into the Montana Natural Heritage
Program Point Observation Database (POD);
copies of the Idaho data have been sent to the
Idaho Conservation Data Center (CDC) in Boise.
We collected voucher specimens of all Species of
Concern (SOC) that we discovered, as well as
representatives of many other non-SOC taxa;
vouchers were preserved in 95% ethanol in order
to permit future genetic analyses. We sent all slug
specimens from Idaho and Montana west of the
Continental Divide to taxonomic experts, and their
identifications have been verified.

Results and Discussion

Overview

We conducted a total of 108 site surveys in 2005
(Appendix B). These were distributed on the
Montana Forests as follows: Beaverhead-
Deerlodge (13), Bitterroot (0), Custer (8), Flathead
(9), Gallatin (17), Helena (13), Kootenai (5), Lewis
& Clark (19), Lolo (1). We distributed surveys on
the Idaho Forests as follows: Clearwater (11),
Idaho Panhandle (4), Nez Perce (8). Some
landscapes within the survey area of the Northern
Region were bypassed because they fell mostly
outside U.S. Forest Service land ownership or they
had received prior survey attention in recent years.
Therefore, we conducted few surveys in the
Lower Salmon River area of Idaho and none in the
Bitterroot Mountains, Mission Mountains, and
Swan Valley of northwestern Montana, despite
these being areas of significant land mollusk
endemism and with additional records of rare
regional species (Solem 1975; Fairbanks 1984;
Frest and Johannes 1995, 1997, 2001; Hendricks
1998,2003).

We documented 83 new locations for ten SOC taxa
and three SOI taxa during our 2005 surveys (Table
3): Selway Forestsnail Allogona lombardii (3
sites), Nimapuna Tigersnail Anguispira nimapuna
(4 sites), Striate Disc Discus shimeki (1 site),

Berry's Mountainsnail Oreohelix strigosa berryi
(6 sites), Robust Lancetooth Haplotrema
vancouverense (4 sites), Humped Coin
Polygyrella polygyrella (6 sites), Fir Pinwheel
Radiodiscus abietum (16 sites), Pale Jumping-slug
Hemphillia camelus (9 sites), Pygmy Slug
Kootenaia burkei (6 sites), Reticulate Taildropper
Prophysaon andersoni (1 site), Smoky
Taildropper Prophysaon humile (16 sites), Lyre
Mantleslug Udosarx lyrata (2 sites), and Sheathed
Slug Zacoleus idahoensis (9 sites). Most new
locations are from west of the Continental Divide in
moist forest habitats (e.g., western redcedar,
western hemlock, grand fir, mesic Douglas-fir).
Two SOC and one SOI slug species, all discovered
on the Kootenai National Forest, are new to the
known mollusk fauna of Montana: Pale Jumping-
slug, Pygmy Slug, and Reticulate Taildropper. The
2005 survey added seven new Montana locations
and nine new Idaho locations for a third SOC slug
species, Smoky Taildropper, which was
documented in Montana only once prior to 2004.
We were most surprised by our failure to document
any locations of Magnum Mantleslug (Magnipelta
mycophaga) during our autumn 2005 survey,
although several new western Montana sites were
found during other surveys in 2004 and spring 2005
(e.g., Appendix E; Hendricks 2005).

Table 3. Number of survey sites where Species-of-Concern land mollusks were detected on Northern Region Forests during
the 2005 survey (n = 108 sites). G Ranks of species in bold changed as a result of our 2005 surveys (see text for previous
ranks). Species common names with an * are newly documented Montana fauna.

Montana 3

Idaho"

SPECIES

GRANK

B-D

BI

CU

FL

GA

HE

KO

L-C

LO

CL

I-P

N-P

SNAILS

Selway Forestsnail

Allogona lombardii (ID)

Gl

3

Nimapuna Tigersnail

Anguispira nimapuna (ID)

Gl

1

3

Striate Disc

Discus shimekii (MT, ID?)

G5

1

Robust Lancetooth

Haplotrema vancouverense (MT, ID)

G5

2

2

Berry's Mountainsnail

Oreohelix strigosa berryi (MT)

G5T2

1

1

4

Humped Coin

Polygyrella polygyrella (MT, ID)

G3

3

3

Fir Pinwheel

Radiodiscus abietum (MT, ID)

G4

1

4

6

2

3

SLUGS

Pale Jumping-slug*

Hemphillia camelus (MT, ID)

G4

1

2

1

5

Pygmy Slug*

Kootenaia burkei (MT, ID)

G2

4

2

Reticulate Taildropper*

Prophysaon andersoni (MT, ID)

G5

1

Smokey Taildropper

Prophysaon humile (MT, ID)

G3

5

1

1

6

1

2

Lyre Mantleslug

Udosarx lyrata lyrata (MT, ID)

G2T2

2

Sheathed Slug

Zacoleus idahoensis (MT, ID)

G3G4

5

2

2

a Montana Forests codes: Beaverhead-Deerlodge (B-D), Bitterroot (BI), Custer (CU), Flathead (FL), Gallatin (GA), Helena (HE),

Kootenai (KO), Lewis & Clark (L-C), Lolo (LO).

b Idaho Forest codes: Clearwater (CL), Idaho Panhandle (I-P), Nez Perce (N-P).

As a result of the 2005 surveys, global ranks
shifted downward for five species. The Global
Rank of Humped Coin changed from G2G3 to G3
(see Appendix A for definitions). The Fir Pinwheel
changed from G3 to G4, Pale Jumping-slug
changed from G3G4 to G4. The Pygmy Slug
changed from G1G2 to G2. The Smoky
Taildropper changed from G2 to G3. In summary,
the 2005 Northern Region survey made a
significant contribution to our understanding of the
current status of several land mollusk species of
conservation interest.

At least some SOI G4-G5 taxa we found during
our 2005 surveys may prove to be distinct from
related coastal populations, as their disjunct
distributions are similar to some vertebrate
amphibian taxa (e.g., Dicarnptodon, Ascaphus,
Plethodon) now split into coastal and Rocky
Mountain sister species. Thus, we think it desirable
to conduct genetic analyses of several mollusk
SOC and SOI taxa to determine if they represent
forms that merit full species status. These will be
identified in the species accounts that follow of
SOC/SOI taxa encountered during the 2005 survey.

Finally, we recorded 21 additional terrestrial
mollusk species (including exotics) as we
encountered them during our 2005 surveys. These
species are not currently recognized as SOC or
SOI, nor are they considered to merit such status,
and will not be discussed further in this report.
Distribution maps showing locations where we
found these taxa are in Appendix F (native Non-
SOC) and Appendix G (exotic species). Not
included in Appendix F is a map showing our two
2005 records of the Chrome Ambersnail (Catinella
rehderi) from Carbon and Fergus counties,
Montana. Species of Catinella are impossible to
identify to species based on shells alone (T. Pearce
personal communication), so our identification of
shells from these sites remains tentative, and
influenced by one prior Montana record from
Meagher County (Pilsbry 1948).

Species Accounts

Selway Forestsnail (Allogona lombardii): We
found this species at three Idaho sites between
1600-3700 ft elevation, all in Idaho County

(Appendix B and E). The Selway Forestsnail is an
Idaho endemic known only from the Lochsa-
Selway and Lower Salmon corridors of Idaho
County (Smith 1943; Frest and Johannes 1997,
2001). Canopy at the 2005 sites included grand fir,
western hemlock, and alder; as many as 1 1
individuals were found on or under bryophyte mats.

Nimapuna Tigersnail (Anguispira nimapuna): We
found this species at four Idaho sites between
1500-2550 ft elevation, all in Idaho County
(Appendix B and E). The Nimapuna Tigersnail is
an Idaho endemic known only from the Clearwater,
Lochsa, and Selway rivers drainages (Baker 1932;
Smith 1943; Frest and Johannes 1995, 2001).
Canopy at the 2005 sites included western
redcedar and grand fir, with some alder, paper
birch, Douglas-fir and/or ponderosa pine; as many
as 10 live animals were found under wood or on
bryophyte mats among dense ferns.

Striate Disc {Discus shimeki): We found this
species at one Montana site in Gallatin County, at
5832 ft elevation (Appendix B). The Striate Disc
has a wide distribution in western North America
(Pilsbry 1948; Frest and Johannes 1993; Forsyth
2004) and is ranked G5. It is a Montana SOC
because there are less than 10 documented
occurrences in the state (see map in Appendix E).
Canopy at the 2005 site included Douglas-fir and
subalpine fir; two shells were present on a
bryophyte mat.

Robust Lancetooth (Haplotrema vancouverense):
We found this species at four Idaho sites between
2100-3350 ft elevation, in Bonner, Clearwater, and
Shoshone counties (Appendix B and E). The
Robust Lancetooth has a wide distribution in the
Pacific Coast states and British Columbia (Forsyth
2004) and is ranked G5. It is a new Montana SOC
because of only two historical records from
Sanders County (Brunson and Osher 1957) and no
new records in recent years. Frest and Johannes
(2001) listed the Robust Lancetooth as rare and
only from northern Idaho. Populations in northern
Idaho and northwestern Montana appear disjunct
from the main coastal range, and should be
examined genetically to determine if they actually
are sister species. Canopy at the 2005 sites

included western redcedar, grand fir, western
hemlock, alder, or paper birch; live individuals and
shells were found under wood or bryophyte mats.

Berry's Mountainsnail (Oreohelix strigosa
berry i): We found this subspecies at six sites in
central Montana between 4250-6400 ft elevation, in
Broadwater, Carbon, Fergus, and Meagher
counties (Appendix B and E). Berry's
Mountainsnail is a narrowly distributed subspecies,
largely restricted to central Montana and the Black
Hills (Frest and Johannes 1993), of a species that is
found throughout western North America (Pilsbry
1939; Forsyth 2004). It is most abundant in the
island mountain ranges of central Montana,
especially the Big Snowy Mountains (Berry 1916).
Canopy at the 2005 sites included aspen, Douglas-
fir, Engelmann spruce, subalpine fir, cottonwood
and maple. Few live animals were found, but
hundreds of shells were scattered on and under
limestone talus and soil, bryophyte mats, and leaf
litter.

Humped Coin {Polygyrella polygyrella): We
found this species at six Idaho sites between 1500-
3600 ft elevation, in Clearwater and Idaho counties
(Appendix B and E). The Humped Coin, first
described from Montana and Idaho by Bland and
Cooper (1861) and Cooper (1868), is also present
in adjacent Washington and Oregon (Frest and
Johannes 1995, 2001). In 2005, we found this
species only in the Clearwater and Lochsa river
drainages, but it is present farther north in Idaho,
and all known Montana sites are clustered in
Sanders and Mineral counties (Hendricks 2003,
2005). Canopy at the 2005 sites included western
redcedar, western hemlock, grand fir, alder, black
cottonwood, and mountain maple. Live animals
were found at all sites, with as many as 44 found
on ferns and bryophyte mats.

Fir Pinwheel (Radiodiscus abietum): We found
this species at 11 Idaho sites between 1500-5500 ft
elevation, in Bonner, Clearwater, Idaho, and
Shoshone counties, and at five sites in Montana
between 2200-3450 ft elevation, in Flathead,
Lincoln, and Sanders counties (Appendix B and E).
The Fir Pinwheel is restricted to northern Idaho,
western Montana, and adjacent parts of Oregon

and Washington (Brunson and Russell 1967; Frest
and Johannes 1995, 2001; Hendricks 2003, 2005).
Canopy at the 2005 sites included western
redcedar, grand fir, Douglas-fir, western hemlock,
subalpine fir, alder, water birch, cottonwood,
western larch, and Pacific yew. Up to 11 live
individuals were present at 15 sites where they
were detected, mostly under downed wood, but
also rocks and bryophyte mats. At the remaining
site only a single shell was detected.

Pale Jumping- slug (Hemphillia camelus): We
found this species at eight Idaho sites between
1600-5 100 ft elevation, in Clearwater, Idaho, and
Shoshone counties, and at one site in Montana at
2840 ft elevation, in Lincoln County (Appendix B
and E). This is the first Montana record for this
species (Frest and Johannes 1995; Hendricks 2003)
(see Appendix E; Pilsbry 1948; Hendricks 2003).
The Pale Jumping-slug appears to be restricted to
northern Idaho, and adjacent parts of Washington,
British Columbia, Alberta, and now Montana (Frest
and Johannes 1995, 2001; Forsyth 2004). Frest and
Johannes (1997, 2001) suggested that individuals
from the Lower Salmon River drainage might
represent a taxon distinct from that to the north, but
this possibility has not been resolved. Canopy at
the 2005 sites included western redcedar, grand fir,
subalpine fir, Engelmann spruce, western hemlock,
Pacific yew, alder, and birch. Up to six individuals
were found on and under downed wood, and on
bryophyte mats among ferns and dogwood.

Pygmy Slug (Kootenaia burkei): We found this
species at two Idaho sites between 2600-2700 ft
elevation, in Bonner and Boundary counties, and at
four sites in Montana between 2200-2850 ft
elevation, in Lincoln and Sanders counties
(Appendix B and E). These are the first Montana
records for this species, and more than double the
number of total known localities. Only recently
was this species described from animals discovered
at five Idaho sites. (Leonard et al. 2003). Canopy
at the 2005 sites included western redcedar,
western hemlock, grand fir, Pacific yew, Douglas-
fir, paper birch, alder, black cottonwood, western
larch, and western white pine. Up to four
individuals were found on and under downed wood
and bark among leaf litter, and on bryophyte mats.

Reticulate Taildropper {Prophysaon andersoni):
We found this species at one site in Montana at
2200 ft elevation, in Sanders County (Appendix B
and E). This is the first Montana record for this
species. It has rarely been found in northern Idaho
(B. Leonard personal communication), and Frest
and Johannes (2001) thought it might not be present
at all in northern Idaho, despite the tentative
records of Smith (1943). This species is
widespread in coastal British Columbia,
Washington, Oregon, and northern California
(Forsyth 2004). Populations in northern Idaho and
northwestern Montana appear disjunct from the
main coastal range, and should be examined
genetically to determine if they actually are sister
species. Idaho populations of the congeneric Blue-
gray Taildropper (P. coeruleum) and Papillose
Taildropper (P. dubium) also appear disjunct from
the coastal populations (Leonard et al. 2003;
Ovaska et al. 2004), and these too deserve genetic
comparison to determine their species status; both
species are currently ranked G4, and the Reticulate
Taildropper is ranked G5 (Table 1). Canopy at the
2005 site included western redcedar, grand fir,
black cottonwood, paper birch, and Pacific yew.
Two individuals were found under downed wood.

Smoky Taildropper {Prophysaon humile): We
found this species at nine sites in Idaho between
1500-4250 ft elevation, in Bonner, Clearwater, and
Idaho counties, and at seven sites in Montana
between 2800-4150 ft elevation, in Flathead,
Lincoln, and Missoula counties (Appendix B and
E). This species is known only from northern
Idaho and adjacent northwestern Montana (Pilsbry

1948; Frest and Johannes 1995, 2001; Hendricks
2005). Canopy at the 2005 sites included western
redcedar, grand fir, Douglas-fir, Engelmann spruce,
subalpine fir, lodgepole pine, western hemlock,
alder, paper birch, and cottonwood. Up to nine
individuals were found mostly under downed wood,
bryophyte mats, or rocks.

Lyre Mantleslug (Udosarx lyrata): We found this
species at two Idaho sites between 2100-3600 ft
elevation, in Clearwater and Idaho counties
(Appendix B and E). This species is restricted to
northern Idaho and adjacent parts of western
Montana (Webb 1959; Russell and Webb 1980;
Frest and Johannes 1995, 2001; Hendricks 2003).
Two subspecies are described, but we are unable
to distinguish these and assign our records only to
the species level. Canopy at the 2005 sites
included western redcedar, grand fir, and alder. Up
to four individuals were found under downed wood
and wet bark.

Sheathed Slug (Zacoleus idahoensis): We found
this species at nine Idaho sites between 2000-5500
ft elevation, in Bonner, Clearwater, and Idaho
counties (Appendix B and E). This species is
restricted to northern Idaho and adjacent
northwestern Montana (Pilsbry 1948; Frest and
Johannes 1995, 2001; Hendricks 2003). Canopy at
the 2005 sites included western redcedar, grand fir,
western hemlock, Douglas-fir, Engelmann spruce,
subalpine fir, alder, and paper birch. Up to 1 1
individuals were found under wet downed wood or
on bryophyte mats.

Conclusions and Recommendations

The number of new locations we discovered in
2005 for land mollusk species of conservation
concern in the USFS Northern Region area
underscores our conclusion that current knowledge
of the distribution, ecology, and status of this suite
of species is woefully inadequate and largely
fragmentary. We think at least one additional year
of non-random surveys, similar to those of 2005, is
needed to fill distribution gaps and gather additional
habitat information. We also feel that a minimum
of two additional years of random site surveys
(beyond the one of non-random surveys) are
needed in order to fully document species
distributions and habitat associations and to
determine site occupancy rates, while addressing
detection probabilities. This will provide a measure
of status in various habitats. Developing predictive
habitat models may also prove useful for some
species groups, especially those associated with the
moist forest types mentioned earlier. Recent
examples of the use of predictive models for
conservation management of rare terrestrial
mollusks in the Pacific Northwest are Dunk et al.
(2004) and Gaines et al. (2005).

Other recommendations include the following: (1)
Survey and modeling efforts should be coordinated
with the Idaho CDC, as well as MTNHP, as the
Idaho CDC expertise with the Idaho land mollusk

fauna exceeds ours; (2) There is a need for genetic
studies to inform current taxonomy of some
species. We think some taxa currently considered
conspecific with coastal populations (e.g., Robust
Lancetooth, Reticulate Taildropper, Blue-gray
Taildropper, and Papillose Taildropper) may prove
to be distinct sister species (see discussions in
Leonard et al. 2003, Ovaska et al. 2004), similar to
the results of recent genetic studies of some
Pacific Northwest amphibian genera (e.g.,
Ascaphus, Dicarnptodon, Plethodon); (3) Finally,
we think it would be useful to conduct some
workshops on land mollusk identification and
management. This will heighten awareness of this
overlooked and poorly understood group of animals,
and provide biologists and managers some of the
basic tools they need to make informed
management decisions.

Besides producing this summary document for the
2005 inventory, we anticipate developing an
illustrated field guide and/or poster that will aid
District Biologists in future survey work they
conduct targeting SOC and SOI land mollusks;
similar information and illustrations for Montana
species will be made available in the near future in
the Montana Natural Heritage Program on-line
Animal Field Guide.

References Cited

Baker, H. B. 1932. New land snails from Idaho
and eastern Oregon. The Nautilus 45:82-
87.

Berry, S. S. 1916. Notes on mollusca of central
Montana. The Nautilus 29: 124-128.

Bland, T., and J. G. Cooper. 1861. Notice of land
and freshwater shells collected by Dr. J. G.
Cooper in the Rocky Mountains, etc., in
1860. Annals of the Lyceum of Natural
History of New York 7:362-370.

Brunson, R. B., and U. Osher. 1957. Haplotrema
from western Montana. The Nautilus
70:121-123.

Brunson, R. B., and R. H. Russell. 1967.

Radiodiscus, new to molluscan fauna of
Montana. The Nautilus 81:1 8-22.

CFR (Code of Federal Regulations). 1985. 36

Code of Federal Regulations. Chapter II
219.19:64.

Cooper, J. G 1868. The shells of Montana.
American Naturalist 2:486-487.

Dunk, J. R., W. J. Zielinski, and H. K. Preisler.
2004. Predicting the occurrence of rare
mollusks in northern California forests.
Ecological Applications 14:71 3-729.

Fairbanks, H. L. 1984. A new species of
Oreohelix (Gastropoda: Pulmonata:
Oreohelicidae) from the Seven Devils
Mountains, Idaho. Proceedings of the
Biological Society of Washington 97: 179-
185.

Forsyth, R. G 2004. Land snails of British
Columbia. Royal British Columbia
Museum, Victoria, BC 188 pp.

Frest, T. J., and E. J. Johannes. 1993. Land snail
survey of the Black Hills National Forest,
South Dakota and Wyoming. Final report
to USDA Forest Service, Black Hills
National Forest and USDI Fish & Wildlife
Service, South Dakota State Office. 156
pp. + appendices.

Frest, T J., and E. J. Johannes. 1995. Interior

Columbia Basin mollusk species of special
concern. Final Report to Interior Columbia
Basin Ecosystem Management Project.
Deixis Consultants, Seattle. 274 pp.

Frest, T. J., and E. J. Johannes. 1997. Land snail
survey of the lower Salmon River drainage,
Idaho. Idaho Bureau of Land
Management Technical Bulletin No. 97-18.

Frest, T. J., and E. J. Johannes. 2001. An

annotated checklist of Idaho land and
freshwater mollusks. Journal of the Idaho
Academy of Science 36:1-51.

Gaines, W. L., A. L. Lyons, and A. Sprague. 2005.
Predicting the occurrence of a rare mollusk
in the dry forests of north-central
Washington. Northwest Science 79:99-
105.

Hendricks, P. 1998. Rediscovery of Discus
brunsoni Berry, 1955 and Oreohelix
alpina (Elrod, 1901) in the Mission
Mountains, Montana, with comments on
Oreohelix elrodi (Pilsbry, 1900). The
Nautilus 112:58-62.

Hendricks, P. 2003. Status and conservation
management of terrestrial mollusks of
Special Concern in Montana. Report to
Region 1, U.S. Forest Service. Montana
Natural Heritage Program, Helena, MT.
67 pp. + appendices.

10

Hendricks, P., Compiler. 2005. Surveys for Animal
Species of Concern in northwestern
Montana. Report to Montana Department
of Fish, Wildlife, and Parks, State Wildlife
Grants Program, Helena, Montana.
Montana Natural Heritage Program,
Helena, MT. 53 pp.

Leonard, W. P., L. Chichester, J. Baugh, and T.
Wilke. 2003. Kootenaia burkei, a new
genus and species of slug from northern
Idaho, United States (Gastropoda:
Pulmonata: Arionidae). Zootaxa 355: 1-16.

Smith, A. G 1943. Mollusks of the Clearwater
Mountains, Idaho. Proceedings of the
California Academy of Sciences, fourth
series, 23:537-554.

Webb, G R. 1959. Two new north-western slugs,
Udosarx lyrata and Gliabates oregonia.
Gastropodia l(3):22-23, 28.

Leonard, W. P., L. Chichester, and K. Ovaska.
2003. Prophysaon dubium Cockerell,
1890, the papillose taildropper (Gastropoda:
Arionidae): distribution and anatomy. The
Nautilus 117:62-67.

Ovaska, K., W. P. Leonard, L. Chichester, T. E.
Burke, L. Sopuck, and J. Baugh. 2004.
Prophysaon coeruleum Cockerell, 1890,
blue-gray taildropper (Gastropoda:
Arionidae): new distributional records and
reproductive anatomy. Western North
American Naturalist 64:538-543.

Pilsbry, H.A. 1939. Land mollusca of North

America (north of Mexico), Volume I Part

1. The Academy of Natural Sciences of
Philadelphia Monographs Number 3 (1): 1-

573.

Pilsbry, H.A. 1948. Land mollusca of North

America (north of Mexico), Volume II Part

2. The Academy of Natural Sciences of
Philadelphia Monographs Number 3
(2):521-1113.

Russell, R. H., and G R.Webb. 1980. The slug
Udosarx lyrata: additional data on
distribution, anatomy, and taxonomy.
Gastropodia 2:8-10.

Solem, A. 1975. Notes on Salmon River Valley
oreohelicid land snails, with description of
Oreohelix waltoni. The Veliger 18:16-30.

11

Appendix A. Global/State Rank Definitions

Heritage Program Ranks

The international network of Natural Heritage Programs employs a standardized ranking system to denote
global (range-wide) and state status. Species are assigned numeric ranks ranging from 1 to 5, reflecting
the relative degree to which they are "at-risk". Rank definitions are given below. A number of factors are
considered in assigning ranks — the number, size and distribution of known "occurrences" or populations,
population trends (if known), habitat sensitivity, and threat. Factors in a species' life history that make it
especially vulnerable are also considered (e.g., dependence on a specific pollinator).

Global Rank Definitions (NatureServe 2003)

G 1 Critically imperiled because of extreme rarity and/or other factors making it highly

vulnerable to extinction
G2 Imperiled because of rarity and/or other factors making it vulnerable to extinction

G3 Vulnerable because of rarity or restricted range and/or other factors, even though it may

be abundant at some of its locations
G4 Apparently secure, though it may be quite rare in parts of its range, especially at the

periphery
G5 Demonstrably secure, though it may be quite rare in parts of its range, especially at the

periphery
Tl-5 Infraspecific Taxon (trinomial) — The status of infraspecific taxa (subspecies or

varieties) are indicated by a "T-rank" following the species' global rank

State Rank Definitions

51 At high risk because of extremely limited and potentially declining numbers,
extent and/or habitat, making it highly vulnerable to extirpation in the state

52 At risk because of very limited and potentially declining numbers, extent and/or
habitat, making it vulnerable to extirpation in the state

53 Potentially at risk because of limited and potentially declining numbers, extent
and/or habitat, even though it may be abundant in some areas

54 Uncommon but not rare (although it may be rare in parts of its range), and usually
widespread. Apparently not vulnerable in most of its range, but possibly cause for
long-term concern

55 Common, widespread, and abundant (although it may be rare in parts of its
range). Not vulnerable in most of its range

Combination Ranks

G#G# or S#S# Range Rank — A numeric range rank (e.g., G2G3) used to indicate uncertainty about
the exact status of a taxon

Qualifiers

NR Not ranked

Q Questionable taxonomy that may reduce conservation priority — Distinctiveness of

this entity as a taxon at the current level is questionable; resolution of this uncertainty may
result in change from a species to a subspecies or hybrid, or inclusion of this taxon in
another taxon, with the resulting taxon having a lower-priority (numerically higher)
conservation status rank

Appendix A - 1

X Presumed Extinct — Species believed to be extinct throughout its range. Not located

despite intensive searches of historical sites and other appropriate habitat, and virtually no
likelihood that it will be rediscovered

H Possibly Extinct — Species known from only historical occurrences, but may neverthe-

less still be extant; further searching needed

U Unrankable — Species currently unrankable due to lack of information or due to substan-

tially conflicting information about status or trends

HYB Hybrid — Entity not ranked because it represents an interspecific hybrid and not a species

? Inexact Numeric Rank — Denotes inexact numeric rank

C Captive or Cultivated Only — Species at present is extant only in captivity or cultivation,

or as a reintroduced population not yet established

A Accidental — Species is accidental or casual in Montana, in other words, infrequent and

outside usual range. Includes species (usually birds or butterflies) recorded once or only a
few times at a location. A few of these species may have bred on the one or two occa-
sions they were recorded

Z Zero Occurrences — Species is present but lacking practical conservation concern in

Montana because there are no definable occurrences, although the taxon is native and
appears regularly in Montana

P Potential — Potential that species occurs in Montana but no extant or historic occurrences

are accepted

R Reported — Species reported in Montana but without a basis for either accepting or

rejecting the report, or the report not yet reviewed locally. Some of these are very recent
discoveries for which the program has not yet received first-hand information; others are
old, obscure reports

SYN Synonym — Species reported as occurring in Montana, but the Montana Natural Heritage

Program does not recognize the taxon; therefore the species is not assigned a rank

* A rank has been assigned and is under review. Contact the Montana Natural Heritage

Program for assigned rank

B Breeding — Rank refers to the breeding population of the species in Montana

N Nonbreeding — Rank refers to the non-breeding population of the species in Montana

Appendix A - 2

Appendix B. USFS Region 1 Survey Sites for Land Mollusks

in 2005

t

bo

Forest 3

State

County

UTM

NAD 27

Site Name

Elev

(ft)

Date

SOC/SOI Taxa b

B-D

MT

Madison

12: 433652E 5032978N

Daisy Creek

6221

6 Oct

B-D

MT

Madison

12: 433327E 5033290N

Daisy Creek

6387

6 Oct

B-D

MT

Madison

12:431197E5042579N

N Meadow Creek

6622

6 Oct

B-D

MT

Madison

12:431390E5051347N

Willow Creek

5723

6 Oct

B-D

MT

Madison

12: 456430E 5000257N

Sphinx Mountain C.G.

6195

7 Oct

B-D

MT

Madison

12:456481E4993881N

Indian Creek

6210

7 Oct

B-D

MT

Madison

12:455242E4971846N

Papoose Creek trail

6311

7 Oct

B-D

MT

Madison

12:453811E4970009N

Gazette Creek

6000

7 Oct

B-D

MT

Beaverhead

12:334379E5051791N

Lacy Creek

6693

20 Oct

B-D

MT

Beaverhead

12:353401E5032771N

Birch Creek Rd

6950

20 Oct

B-D

MT

Madison

12:421598E4967721N

Basin Creek, Ruby River Rd

6946

21 Oct

B-D

MT

Madison

12:421796E4974062N

opposite Dos Creek

6723

21 Oct

B-D

MT

Madison

12: 422342E 4976048N

Dry Fawn Creek

6646

21 Oct

B-D

MT

Madison

12:423738E4981384N

Short Creek

6353

21 Oct

CL

ID

Clearwater

H:644430E5188837N

N. Fork Clearwater River

3540

22 Sep

Raab

CL

ID

Clearwater

1L643771E5187848N

Deception Gulch

3700

22 Sep

Raab

CL

ID

Clearwater

1L630784E5175677N

N. Fork Clearwater River

2745

22 Sep

Popo, Prhu

CL

ID

Clearwater

1L616029E5181048N

N. Fork Clearwater River

2000

22 Sep

Heca, Prhu, Raab,
Zaid

CL

ID

Clearwater

1L618442E5189072N

Skull Creek

2125

23 Sep

Hava, Prhu, Udly

CL

ID

Clearwater

1L610464E5173214N

Washington Creek
Campground

2100

23 Sep

Hava, Prhu, Raab,
Zaid

CL

ID

Clearwater

1L606286E5109880N

Wild Goose Campground

1500

29 Sep

Anni, Popo, Prhu,
Raab

CL

ID

Clearwater

1L684222E5163577N

Lolo Pass

4570

30 Sep

Heca

CL

ID

Clearwater

1L678297E5156289N

DeVoto Memorial Grove

3590

30 Sep

Popo, Prhu, Udly,
Zaid

CL

ID

Clearwater

1L685846E5164927N

Pack Creek, Lolo Pass

5160

30 Sep

Zaid

CL

ID

Clearwater

1L691799E5166331N

Cascade above Brushy Fork

5500

6 Oct

Raab, Zaid

CU

MT

Carbon

12: 633364E 5003082N

Palisades Campground

6312

5 Oct

CU

MT

Carbon

12: 628574E 5001261N

Washoe Creek

6717

10 Oct

t

bo

Forest 3

State

County

UTM
NAD 27

Site Name

Elev

(ft)

Date

SOC/SOI Taxa b

CU

MT

Carbon

12: 632079E 4993907N

Ratine Campground

6388

11 Oct

Osbe

cu

MT

Carbon

12: 623046E 4987040N

Rock Creek road

7552

11 Oct

CU

MT

Carbon

12: 698445E 5006938N

Trappers Cabin Spring

6674

11 Oct

cu

MT

Carbon

12: 693394E 5009266N

Sage Creek

5732

11 Oct

cu

MT

Carbon

12:701129E5005597N

Crooked Creek

7171

12 Oct

Care

cu

MT

Carbon

12: 700432E 5006043N

Tie Flat

7087

12 Oct

FL

MT

Flathead

12: 302229E 5323477N

N. Fork Clark Creek

3820

20 Oct

Prhu

FL

MT

Flathead

12:292962E5333194N

Graves Bay

3750

20 Oct

Prhu

FL

MT

Flathead

11:719845E5361343N

Whelp Creek, at Lion Lake

3440

20 Oct

Prhu

FL

MT

Flathead

12: 306742E 5349804N

Walton

3780

21 Oct

FL

MT

Flathead

12:285169E5374870N

Kootenai Creek

3440

21 Oct

Prhu, Raab

FL

MT

Flathead

11:714612E5363026N

Badrock Canyon

3050

21 Oct

Prhu

FL

MT

Flathead

1L712272E5375118N

Glacier Rim river access

3180

25 Oct

FL

MT

Flathead

1L697962E5382199N

Big Creek

4140

25 Oct

FL

MT

Flathead

1L688922E5409241N

Red Meadow Creek

4240

26 Oct

GA

MT

Meagher

12:536538E5119371N

Smith Creek

6257

1 Oct

GA

MT

Meagher

12:548915E5115237N

Dugout Creek

6971

1 Oct

GA

MT

Meagher

12:550001E5114296N

Lodgepole Creek

6700

1 Oct

GA

MT

Meagher

12:543965E5112874N

Deep Creek

6456

1 Oct

GA

MT

Meagher

12:544502E5124436N

Castle Creek

6039

lOct

GA

MT

Park

12:537684E5038151N

Pine Creek trailhead

5668

4 Oct

GA

MT

Park

12:537107E5014922N

Mill Creek

5688

5 Oct

GA

MT

Sweetgrass

12: 562854E 5033683N

Aspen Campground

5407

12 Oct

GA

MT

Sweetgrass

12:561089E5037507N

Falls Creek Campground

5242

12 Oct

GA

MT

Gallatin

12:482099E5013744N

Dudley Creek, along Hwy 191

5946

13 Oct

GA

MT

Gallatin

12: 483550E 5015692N

Goose Creek

5987

13 Oct

GA

MT

Gallatin

12:484959E5031594N

Spire Rock Campground

5832

13 Oct

Dish

GA

MT

Gallatin

12: 499497E 5042474N

Moser Creek

6310

13 Oct

GA

MT

Gallatin

12: 503346E 5090109N

road to Flathead Pass

6323

14 Oct

GA

MT

Gallatin

12: 506564E 5084007N

Fairy Creek

6549

14 Oct

GA

MT

Gallatin

12:508149E5081951N

Cache Creek

6192

14 Oct

t

bo

Forest 3

State

County

UTM
NAD 27

Site Name

Elev
(ft)

Date

SOC/SOI Taxa b

HE

MT

Meagher

12:492933E5129289N

Battle Mountain Road

6380

19 Sep

HE

MT

Broadwater

12:488119E5132001N

Deep Creek

5080

19 Sep

HE

MT

Broadwater

12:457104E5164506N

Avalanche Creek

4268

19 Sep

Osbe

HE

MT

Jefferson

12:435798E5147472N

Crystal Creek

5224

27 Sep

HE

MT

Jefferson

12:428271E5141122N

Warm Springs Creek

4680

27 Sep

HE

MT

Broadwater

12:446064E5142057N

Weasel Creek

5390

27 Sep

HE

MT

Lewis & Clark

12:449455E5187615N

Beaver Creek

6274

30 Sep

HE

MT

Lewis & Clark

12:446328E5189656N

Beaver Creek

4809

30 Sep

HE

MT

Lewis & Clark

12:440972E5177782N

Nelson Road

4311

30 Sep

HE

MT

Lewis & Clark

12:386784E5180761N

S. Fork Little Prickly Pear Cr.

5240

5 Oct

HE

MT

Lewis & Clark

12:387145E5181566N

Bear Trap Gulch

5100

5 Oct

HE

MT

Lewis & Clark

12:381909E5192006N

Poorman Creek

5320

5 Oct

HE

MT

Lewis & Clark

12:379065E5192491N

Poorman Creek

5000

5 Oct

I-P

ID

Bonner

11:565950E5341771N

Lightening Creek

2610

27 Oct

Hava, Kobu, Prhu,
Raab, Zaid

I-P

ID

Shoshone

11:588051E5284451N

West Fork Eagle Creek
(Settlers Cedar Grove)

3320

27 Oct

Hava, Heca, Raab

I-P

ID

Boundary

11:562646E5407425N

Meadow Creek Campground

2310

4 Nov

Zaid

I-P

ID

Boundary

11:541530E5390916N

Snow Creek

2685

4 Nov

Kobu

KO

MT

Lincoln

11:624604E5398084N

Peck Gulch and Landing

2480

26 Oct

KO

MT

Lincoln

11:580694E5339821N

Ross Creek Cedar Grove

2840

26 Oct

Kobu, Prhu, Raab

KO

MT

Lincoln

11:582176E5388329N

Yaak Falls Campground

2400

3 Nov

Kobu, Raab

KO

MT

Lincoln

11:579587E5401244N

Red Top Campground

2840

3 Nov

Heca, Kobu, Raab

KO

MT

Sanders

11:580446E5324068N

Big Eddy Campground

2200

4 Nov

Kobu, Pran, Raab

L-C

MT

Meagher

12:543825E5128430N

Deer Creek

6174

29 Sep

L-C

MT

Chouteau

12: 531512E5257173N

N. Fork Highwood Creek

4577

29 Sep

L-C

MT

Cascade

12:514564E5215409N

Dry Fork

4680

29 Sep

L-C

MT

Meagher

12:543825E5128430N

tributary to Cottonwood Creek

6174

1 Oct

L-C

MT

Golden Valley

12:651613E5177625N

Minerva Creek

4910

2 Oct

L-C

MT

Fergus

12:648557E5179128N

Pole Creek

4955

2 Oct

Care, Osbe

L-C

MT

Fergus

12: 613571E5183942N

Crystal Lake Campground

6109

2 Oct

Osbe

t

bo

Forest 3

State

County

UTM

NAD 27

Site Name

Elev

(ft)

Date

SOC/SOI Taxa b

L-C

MT

Fergus

12:613661E5189516N

Big Rock Creek

5204

2 Oct

Osbe

L-C

MT

Meagher

12:540971E5162557N

Spring Creek

5431

4 Oct

L-C

MT

Meagher

12:541887E5155849N

Spring Creek

5195

4 Oct

Osbe

L-C

MT

Meagher

12:508385E5184971N

Moose Creek

5707

4 Oct

L-C

MT

Meagher

12:520377E5154218N

Fourmile Creek

5850

4 Oct

L-C

MT

Meagher

12:523595E5152921N

headwaters of Fourmile Creek

6371

4 Oct

L-C

MT

Meagher

12:531387E5155015N

headwaters of Brooks Creek

5987

4 Oct

L-C

MT

Teton

12: 372728E 530495 IN

Clary Coulee, near Wind Mtn

5160

6 Oct

L-C

MT

Teton

12: 371077E 5306593N

N. Fork Teton River

5200

6 Oct

L-C

MT

Lewis & Clark

12: 368275E 5273593N

confluence of Beaver Creek
and N. Fork Sun River

4580

6 Oct

L-C

MT

Lewis & Clark

12: 370616E 5274776N

N. Fork Sun River C.G.

4480

6 Oct

LO

MT

Missoula

12:276255E5199886N

Woods Gulch

4150

5 Oct

Prhu

N-P

ID

Idaho

1L565439E5037059N

Allison Creek

3640

27 Sep

Alio, Zaid

N-P

ID

Idaho

1L571344E5078279N

Fish Creek Meadow
Campground

5060

27 Sep

Heca

N-P

ID

Idaho

1L582828E5075274N

Mill Creek

2370

28 Sep

Alio, Popo

N-P

ID

Idaho

1L586708E5074957N

Johns Creek trailhead

2525

28 Sep

Anni, Heca, Popo,
Prhu, Raab, Zaid

N-P

ID

Idaho

1L601568E5072831N

Rainy Day Creek

3445

28 Sep

Heca

N-P

ID

Idaho

1L620257E5083623N

American River Campground

4250

28 Sep

Heca, Prhu

N-P

ID

Idaho

1L580307E5075362N

Castle Creek

2300

28 Sep

Anni, Raab

N-P

ID

Idaho

1L614631E5103162N

O'Hara Creek

1640

29 Sep

Alio, Anni, Heca,
Popo, Raab

a National Forests: B-D (Beaverhead-Deerlodge), CL (Clearwater), CU (Custer), FL (Flathead), GA (Gallatin), HE (Helena), I-P (Idaho Panhandle), KO
(Kootenai), L-C (Lewis and Clark), LO (Lolo), N-P (Nez Perce).

b SOC/SOI taxa codes: Alio (Selway Forestsnail, Allogona lombardii), Anni (Nimapuna Tigersnail, Anguispira nimapuna), Care (Chrome Ambersnail,
Catinella rehderi), Dish (Striate Disc, Discus shimekii), Hava (Robust Lancetooth, Haplotrema vancouverense), Heca (Pale Jumping-slug, Hemphillia
camelus), Kobu (Pygmy Slug, Kootenai burkei), Osbe (Berry's Mountainsnail, Oreohelix strigosa berryi), Popo (Humped Coin, Polygyrella
polygyrella), Pran (Reticulate Taildropper, Prophysaon andersoni), Prhu (Smokey Taildropper, Prophysaon humile), Raab (Fir Pinwheel, Radiodiscus
abietum), Udly (Lyre Mantleslug, Udosarx lyrata), Zaid (Sheathed Slug, Zacoleus idahoensis).

Appendix C. Standardized Data Sheets

Data Form for Terrestrial Mollusk Site Surveys
Locality Information

Ecoregion:

Sample Block:

Site No:

Locality:

State:

County:

Map
Name:

R

Section
Description:

Owner:

Map
Elevation:

FT

Datum:

UTM

Zone:

UTM
East:

UTM
North:

Habitat Information

Area (M 2 )
Searched:

Date:

Observer(s)

Begin
Time:

End
Time:

Total Person
Minutes of Search:

Percentage of Site Searched:
1-25 26-50 51-75 76-100

Percent
Slope:

Aspect: N NE NW

SE SW

W

Habitat Type:

Spring/Seep Streamside Talus Deciduous Forest Conifer Forest Mixed Forest Shrub/Steppe Grassland
Other

Primary Canopy Species:

Overall Percent Canopy Cover:

1-25 26-50 51-75 76-100

Canopy Species Average DBH (cm):
0-5 5-15 15-30 30-60 >60

Photo Frame Number (s)
/ Description(s):

Weather: Clear Partly Cloudy Overcast

Rain

Snow

Air Temp:

°C

Soil Temp:

°C

Soil Moisture:

Dry Damp Wet Standing Water Snow

Rock Type: Igneous Metamorphic

Note Specific Type (e.g. limestone, granite):

Sedimentary

Habitat
Threats:

Mollusk Species Information

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., H001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., H001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., LC001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., G001 A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Appendix C - 1

Grid

I Scale:

Site Map for Terrestrial Mollusk Site Surveys

■m

"▼" >^

>

J

J.

^1

*D

raw a re

)ugh sk<

jtchoft

he site

labeling

r major

features

> such a

s streair

is, talus

slopes,

habitat

cover t

ypes, et

c.

Be sure to indicate where animals were detected and label the following locations on the map: G = GPS reading,
and P-> = photo locations and directions of photos.
Other Notes:

Appendix C - 2

Site ID (ecoregion, sample block, site number)

Date:

Mollusk Species Information Continued

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., H001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., H001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., LC001A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Species:

Number Alive and/or Dead, Size, and Time at First Detection (e.g., 2 alive & 4 dead x 15mm Diameter or TL @ 10 minutes)

Tissue Number (e.g., G001 A)

Voucher Number
& Description:

Substrate Association (Circle):
under wood under 4-20cm rock fragments under >20cm rock fragments
under bryophyte mat on bryophyte mat in rock fracture Other

Other Species

Information

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Species:
(millipedes etc.)

Time at First
Detection:

Voucher
Number:

Voucher Description / Comments:

Other Notes:

Appendix C - 3

Site Information

Ecoregion: One of the 14 ecoregion sections in Montana or 6 in the Idaho Panhandle.

Sample Block: Identify three digit number of the sampling block (range 001-999).

Site No: Identify three digit number of the site being surveyed within each sampling block (range 001-999).

Locality: Describe the specific geographic location of the site so that the type of site is described and the straight-line air

distance from one or more permanent features on a 7.5-minute (1 :24,000 scale) topographic map records the position of the site

(e.g., Large talus slope 1.5 miles north of Engle Peak, N side of FS Road 225).

State: Use the two-letter abbreviation.

County: Use the full county name.

Map Name: List the name of the USGS 7.5-minute (1 :24,000 scale) topographic quadrangle map.

T: Record the Township number and whether it is north or south.

R: Record the Range number and whether it is east or west.

S: Record the Section number

Section Description: Describe location of the site at the l A of l A section level (e.g., SENE indicates SE corner of NE corner).

Owner: Use abbreviation of the government agency responsible for managing the land you surveyed, (e.g. USFS, BLM). If

private land was surveyed list the owner's full name to indicate that you did not trespass.

Map Elevation: The elevation of the site as indicated by the topographic map in feet (avoid using elevations from a GPS)

Datum: The map datum used (typically NAD 27 if off topographic map or WGS84 if off GPS unit on standard setting).

UTM Zone: Universal Transverse Mercator zone recorded on the topographic map.

UTM East: Universal Transverse Mercator easting coordinate in meters as recorded on the topographic map or GPS receiver.

Be sure to note any major differences between UTM coordinates on the map and those on the GPS receiver.

UTM North: Universal Transverse Mercator northing coordinate in meters as recorded on the topographic map or GPS

receiver. Be sure to note any major differences between UTM coordinates on the map and those on the GPS receiver.

Survey Information

Date: Use MM-DD-YY format (e.g. 05/12/00 for May 12 of 2000).

Observers: List names or initials of individuals involved with survey of this site and circle the name of the recorder.

Begin Time: List the time the survey began in 24-hour format.

End Time: List the time the survey ended in 24-hour format.

Total Person Minutes of Search: Record the total person minutes the site was searched (e.g. if one person surveys for 15

minutes and another surveys for 30 minutes, but takes 5 minutes to measure a specimen the total person minutes is 40 minutes).

Area (M 2 ) Searched: Area in square meters that was surveyed.

Percent of Site Searched: Circle the appropriate category.

Percent Slope: Percent slope of site. Enter range if variable.

Aspect: Circle primary aspect of the site.

Habitat Type: Circle the appropriate habitat type.

Primary Canopy Species: List the major plant species in the canopy (e.g., red cedar, western hemlock, grand fir, ninebark)

Overall Percent Canopy Cover: Circle the appropriate category for total canopy cover.

Canopy Species Average DBH: Circle the appropriate category.

Photo Frame Number(s) / Descriptions: The number of the photo as viewed on the camera's view screen and a description of

the contents of the photograph (e.g., #13 = 1 x Oreohelix strigosa and #14-18 = 5 x habitat). Take photos of all portions of the

site and anything else that may be of interest (e.g., millipedes, potential site threats).

Weather: Circle weather condition during survey.

Air Temp: Record air temperature in °C at chest height in the shade. °C = (°F - 32)/1.8

Soil Temp: Record soil temperature in °C at 10 cm depth. °C = (°F - 32)/1.8

Soil Moisture: Circle the appropriate category.

Rock Type: Circle the appropriate category; note specific type if known.

Habitat Threats: Note impacts from grazing, logging, mining, flooding, road building, weeds, fire, etc.

Species Information

For each species, record the genus name and species, if known. If species cannot be identified in the field, place a

brief description of their morphology here. Record the number alive and dead, and size range for individuals

encountered, and time at first detection for the first individual encountered (e.g., 2x15 mm diameter (shells) or TL =

80-90mm (slugs): @ 10 minutes). Record the tissue number or range of tissue numbers for tissue samples collected

(see tissue collection protocols). Record the preliminary museum voucher specimen number and description for

voucher specimens collected (see voucher specimen collection protocols). Circle the substrate the animal was

associated with at time of detection. Record the presence of other species detected at the site (e.g., millipedes), the

time at first detection, and the voucher number and description of animals collected (see voucher and tissue collection

Protocols). AppendixC-4

Appendix D. Example Species Photographs

Radiodiscus abietum (Fir Pinwheel)

Allogona lombardii (Selway Forestsnail)

Poly gyrella poly gyrella (Humped Coin)

Anguispira nimapuna (NimapunaTigersnail)

Appendix D - 1

Udosarx lyrata (Lyre Mantleslug)

Prophysaon humile (Smoky Taildropper)

Hemphillia camelus (Pale Jumping-slug)

Magnipelta mycophaga (Magnum Mantleslug)

Appendix D - 2

Appendix E. Distribution Maps for SOC/SOI Land
Mollusks on USFS Region 1 Lands

Terrestrial Mollusk Data Overview for Montana and Northern Idaho

t

Data Source

* Existing MTN HP Data

* Date Gathered in 2005

+ Existing Data from Idaho CDC

Selway Forests nail [Altogona lombardii)

t

Data Source

* Existing MTNHPData

• Data G altered i n 2005

+ Existing Data from Idaho CDC

Dry Land Forestsnail [AHogona ptycophora sotida)

t

+ Existing Carta from Idaho CDC

Ni ma puna Tigersnail [Anguispira nimapuna)

t

Data Source

* Existing MTNHPDaia

* Dala Gathered in 20O5

+ Existing Data from Idaho CDC

Salmon Oregon ian {Cryptomastix harfordiana)

t

i

+ Existing Data (torn Idaho CI3C

Mission Creek Oregonian (Cryptomastix magnidentata)

t

i

On

+ Existing Data from Idaho CDC

Oregon ian [Cryptomastix mutiani bfandi)

t

+ Existing Data from Idaho CDC

River of No Return Oregon ian [Cryptomastix mullani clappi)

t

i

+ Existing Darta from Idaho CDC

Kingston Oregonian (Cryptomastix sanbumi)

t

i

+ Existing Data from Idaho CDC

Lake Disc (Discus brunsoni)

t

K4

o

Data Source

* Existing MTN HP Data

* Data Gathered in 2005

Marbled Disc {Discus marmorensis)

t

^
^

+ Existing Data from Idaho CDC

Striate Disc (Discus shimekii)

t

Data Source

* Existing MTN HP Data

* Data Gathered in 2005

Robust Lancetooth [Hapfotrema vancouverense)

Data Source

* Existing MTNHP Data

* Data Gathered in 2005

Salmon Coil (Heticodiscus salmonaceus)

^
-k

+ Existing Data from Idaho CDC

Pale Jump in g-slug {Hemphillia camelus)

t

^
^i

Data Source

* E*i sting MTN H P Data

» Data Gathered in 2005

+ Existing Data from Idaho CDC

Marbled J um ping-slug (Hemphitlia danietsf)

t

on

Data Source

* Existing MTN HP Data

* Data Gathered in 2005

Pygmy Slug {Kootenaia burkei)

t

Data Source

* Existing MTNHPData

■ Data Gathered in 2005

+ Existing Data from Idaho CDC

Magnum Mantle-slug {Magnipelta mycophaga)

t

00

Data Source

* Exi sting M TNH P Data

■ Data Gathered in 2005

+ Existing Data from Idaho CDC

Alpine Mountainsnail {Oreohefix alpina)

t

Data Source

* Existing MTNHP Data

* Data Gathered in 2005

Bitterroot Mountainsnail (Oreohelix amariradtx)

t

Data Source

* Existing MTN HP Data
+ Data Gathered in 2005

Keeled Mountainsnail {Oreohelix carinifera)

t

i

Data Source

* Existing MTN H P Data

* Data Gathered in 2005

Carinate Mountains nail {Oreohetix efrodi)

t

Data Source

* Easting MTN HP Data

• Data Gathered in 2005

VAVr^i

Seven Devils Mountainsnail (Oreohelix hammeri)

+ Exiting Data from Idaho CDC

Costate Mountainsnail (Oreohelix idahoensis)

t

+ Existing Data from Idaho CDC

Deep Slide Mountainsnail (Oreohelix intersum)

A U^H

+ Existing Data from Idaho CDC

Boulder Pile Mountains nail {Oreohetix jugalis)

on

+ Existing Data from Idaho CDC

Berry's Mountains nail {Oreohelix strigosa berryi)

t

Data Source

* Exi sting MTM H P Data
+ Data Gathered in 2005

Striate Mountainsnail {Oreohelix strigosa goniogyra)

t

00

+ Enisling Data from Idaho CDC

Whorl ed Mountainsnail (Oreohetix vortex)

* Existing Data from Idaho CDC

Lava Rock Mountains nail {Oreohelix wattoni)

t

i

+ Existing Data from Idaho CDC

Gallatin Mountainsnail (Oreohetix yavapai mariae)

t

i

Data Source

Existing MTNHP Daia
Data Gathered in 2005

Western Flat-whorl (Planogyra clappi)

+ Existing Data from Idaho CDC

Humped Coin (Potygyrefla polygyrefla)

t

Data Source

* Existing MTN HP Data

• Date Gathered in 2005

+ Existing Data from Idaho COG

Thin lip Tightcoil (Pristiloma idahoense)

t

i

+ Existing Data from Idaho CDC

"lA hT"

Reticulate Taildropper (Prophysaon andersoni)

i

Data Source

* Existing MTNHPData

* Data Gathered in 2005

Smoky Taildropper (Prophysaon humile)

t

i

on

Data Source

* Existing MTNHP Data

* Data Gathered in 2005

4 Existing Data from Idaho CDC

Fir Pin wheel (Radiodiscus abietum)

t

Data Source

* Existing MTNHP Data

* Data Gathered in 2005

+ Existing Data from Idaho CDC

Lyre Mantles lug (Udosanc lyrata)

t

00

Data Source

* Existing MTNHP Data

■ Data Gathered in 2005

+ Existing Data from Idaho CDC

Sheathed Slug {Zacoleus idahoensis)

t

Data Source

* Esi sting MTN H P Data

* Data Gathered in 2005

+ Existing Data from Idaho CDC

Appendix F. Distribution Maps for Non-SOC Land Mollusks

on USFS Region 1 Lands

Idaho Forestsnall {Allogona ptycophora)

t

Data Source

* Existing MTNHP tela

• Data Gathered In 2005

Banded Tigers nail [Anguispira kochi)

t

Data Source

* Existing MTNHP Data

• Daia Gathered in 2005

+ Existing Data from Idaho CDC

Mellow Column {Columella columella)

t

Data Source

* Existing MTNHP Dala

• Data Gathered in 2O05

XA VrJ

Toothless Column [Columella edentuta)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Coeur d'Alene Oregon ian (Cryptomastix mutlani)

t

oi

Data Source

* Existing MTNHP Dsla

• Data Gathered in 2005

IaUI

Forest Disc {Discus whitneyi)

t

on

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Brown Hive {Euconutus fuivus)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Minute Gem (Hawaiia minuscula)

t

Oo

Data Source

* Existing MTNHP Daia

• Data Gathered in 2005

Spruce Snail {Microphysula ingersofli)

t

^o

Data Source

* Existing MTNHP Daia

* Data Gathered in 2005

1-A.vrH

Blue Glass (Nesovttrea binneyana)

K4

o

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Rocky Mountainsnail {Oreohetix strigosa)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Subalpine Mountainsnail {Oreohelix subrudis)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Silky Val Ionia (Vatlonia cyciophoreita)

Data Source

* Existing MTNHP Da\a

• Data Gath&red in 2005

Multirib Vallonia {Vaitonia gracilicosta)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Weste rn G lass -s n a i I ( Vitrina pellucida)

^
^i

Data Source

* Existing MTNHPDala

• Data Gathered in 2O06

lAVrM

Quick Gloss (Zonitoides arboreus)

on

Data Source

* Existing MTNHP Dala

• Data Gathered in 2005

Appendix G. Distribution Maps for Exotic Land Mollusks

on USFS Region 1 Lands

Black Arion (Arion ater)

Data Source

* Existing MTNHP Daia

• Data Gathered in 2005

Dusky Arion {Arion subfuscus)

t

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Meadow Slug [Deroceras taeve)

t

Data Source

* Existing MTNHP Dala

• Data Gathered in 2005

Gray Field slug {Deroceras reticutatum}

Data Source

* Existing MTNHP Data

• Data Gathered in 2005

Giant Gardens lug {Umax maximus)

t

i

Data Source

* Existing MTNHP Dal*

• Data Gathered in 2005